Content gaps – make an inventory of the site’s key content assets, are they lacking any foundational/cornerstone content pieces, non-existent content types, or relevant topic areas that haven’t been covered? What topics or content are missing from your competitors? Can you beat your competitors’ information-rich content assets? Useful guides on Content Gap Analysis:

If you havent see it already, check out the links in shor's comment below - there are some great resources in there. In some cases you can also consider surveying your current audience or customers through email, on-site surveys or SurveyMonkey.  Be sure to ask for some profiling information that you can use for determining specific persona needs like age, sex, location, etc. (Probably best not to make it sound like a creepy text chat like I just did though...)  :)
2. Targeted Keyword Discovery: Ideally you’ll want to do keyword research based on what the audience wants, not solely on what content the site already has (or plans to have sans audience targeting), which may be limited. I can do keyword research on health conditions and drugs (content I have on my site) and determine what the general population is searching for and optimize my current content, or I can cast my net wide and look at what my target audience wants first, then do my keyword research. You may find there are needs that your site is not meeting. Knowing my senior audience is interested in primarily in prescription drug plans and cheap blood pressure medication, I can first make sure I’m providing that content, and then further determine the top keywords in these areas (in the next article Step 2), and use those terms in relevant and high visibility areas on my site.
Stickers are essentially mini-posters, and advertisers have been using them for decades to get the word out without technically breaking the law. They hand them out to teams who then go out and plaster them over public buildings, bus stops and street signs. When the authorities complain, they say “oh, we only gave them to our customers. We have no control over where they put them.”
Practicing SEO now for over a decade, I don’t often come across many blog posts on the subject that introduce me to anything new — especially when it comes to link building. However, I must admit, after reading your article here I had to bookmark it to refer back to it in the future, as I’m sure it will come in handy when doing SEO for my websites later on down the road.
Thanks for bringing up this point - I agree Eric - competitive positioning can help you determine value that you bring to the table that your competitors dont.  I'm all for it.  Neilsen does some reports that provide awareness, likelihood to recommend, sentiment and other insightsfor your site/brand and your competitors. You can also pull some of that type of insight out of social listening platforms like NetBase, SM2, Radian6, Dow Jones, Nielsen, and so many others.  I've even done some hacked compeitove sentiment comprisons before using Search: searching for [brand or feature] + "like", "love", hate", "wish" etc. 
Who doesn’t love quizzes? They are enjoyable, shareable and hard to resist. Design a quiz for your website that somehow relates to your brand. If your company sells jeans, for instance, you could create a quiz called, “What are the best jeans for your body?” and in the results, show the brand’s recommended jeans. But remember—before showing results, be sure to capture the visitor’s e-mail address.
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Hi Brain, I am a young business owner who has had 4 different websites in the last 2 years but none of them were successful as I would have liked due to lack of SEO. Now I am in process of starting another business and I felt it was time for me to learn about SEO myself. I must say the information you have provided is invaluable and extremely helpful!! I am learning on the go and you are my biggest contributor. Thank you Sir!
Guest post on other blogs, and invite other bloggers to guest post on your site. Guest posts are a great way to drive traffic between related blogs, and allow you to gain readers that might not normally make it to your site. Be sure to allow any guest posters to link back to their own site, and share any guest posts the same as you would your own posts.
To gain more customer engagement, the website must reach its visitors/customers efficiently. Obviously, you want the visitors to read your site content. Check the forms and click through on your Call To Actions (CTA’s) when they arrive on your web page. These features initiate user engagement in action, but it is essential to comprehend the in-depth analysis.

Like you I am a scientist and like you did in the past, I am currently working on translating great scientific literature into tips. In my case it’s child development research into play tips for parents. I can already see that the outcome of my experiment is going to be the same as yours. Great content but who cares. I hadn’t even thought about my key influences. I know some important ones, but don’t see how they would share my content. I thought I was writing content for my potential customers. Is your SEO that works course the same as the content that gets results course? Sorry if I sound a bit dim asking that question.
Hey Sammy, I would always advise against buying traffic, social followers, or anything else in that area. It mostly ends up being a vanity metric without business benefits. It’s always better to earn the traffic by creating a valuable, high-quality website and marketing it properly. When you do that, you attract the kind of visitors who are interested in what you have to offer, which is usually better for the bottom line.
It’s free to be active in online groups and on websites that are relevant to your business and community—and it helps you to obtain more traffic. Comment on blogs and social media posts, answer questions people are posting, and participate in conversations about your industry. The more you engage with your community, the more exposure and profile visits you get.
I completely agree that defintion of a target audience isa great first step, but would ask if adding in competitors to the analysis (mentioned here as a later step) helps draw out who your target audience would be via comparisons, i.e. showing who you are an who you are not - would be very interested to hear opinions on how this tactic can be used within the overall step in coordination with targeted keyword discovery.

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